The big bar to reproduction between those with sufficiently different genomes is the different proteins and signal pathways that need to exist. A hybrid often has such a jumbled set of proteins and signal elements that, assuming it can even live, reproduction is almost impossible. That's why humans cannot mate with oak trees but could mate with a close cousin species like Neanderthal.
In the case of Chrom-2 the genes for the proteins and signal elements are all there just in a different location than before the fusion. It helps that in the transcription process the transcriptase RNAs are not specifically targeted to a specific chromosome but to the control regions of the gene.
Edited by AZPaul3, : No reason given.
Edited by AZPaul3, : No reason given.
Edited by AZPaul3, : I quit.