Register | Sign In


Understanding through Discussion


EvC Forum active members: 64 (9164 total)
3 online now:
Newest Member: ChatGPT
Post Volume: Total: 916,788 Year: 4,045/9,624 Month: 916/974 Week: 243/286 Day: 4/46 Hour: 1/0


Thread  Details

Email This Thread
Newer Topic | Older Topic
  
Author Topic:   The End of Evolution By Means of Natural Selection
Percy
Member
Posts: 22492
From: New Hampshire
Joined: 12-23-2000
Member Rating: 4.9


Message 481 of 851 (556819)
04-21-2010 8:59 AM
Reply to: Message 477 by Wounded King
04-21-2010 6:45 AM


Re: Mutations Revisited 2
Wounded King writes:
There are many examples of genes that have been identified as the bases of hybrid sterility and inviability...
How many closely related species are different in only their allele frequencies? Squirrels? Finches? Cats? Any species at all?
Say you have two members of the same species that are mutually unfertile because they each have one of a pair of antagonistic alleles. Now consider these same two individuals except they're in two closely related populations. Nothing has changed genetically, but in one case we consider them the same species and in the other we don't. You're going to explain fine distinctions like this to Faith?
Another related issue is that Faith isn't looking at this at a genetic level. She sees isolation occurring due to physical differences caused by allele combinations that never occurred in the parent population.
The objection to Faith's scenario isn't that it's impossible, because it isn't. The objection is that it's very rare, so rare that there are no known examples of it in the natural world. (I don't actually believe that, but since we've done the equivalent of the human genome project on very few other species I don't expect it can be contradicted at the current time. But even if it is it still must be very rare.) How could it be that we can find no examples of species produced by the process Faith thinks is the cause of all speciation?
--Percy

This message is a reply to:
 Message 477 by Wounded King, posted 04-21-2010 6:45 AM Wounded King has not replied

Replies to this message:
 Message 484 by ZenMonkey, posted 04-21-2010 9:48 AM Percy has seen this message but not replied
 Message 497 by Faith, posted 04-21-2010 5:12 PM Percy has replied

  
Percy
Member
Posts: 22492
From: New Hampshire
Joined: 12-23-2000
Member Rating: 4.9


Message 482 of 851 (556820)
04-21-2010 9:03 AM
Reply to: Message 479 by RAZD
04-21-2010 7:49 AM


Re: Mutations Revisited 2
RAZD writes:
PaulK has his finger on another problem with the Faith Hypothesis: it is not possible for a single organism to carry all the alleles in existence today, they can only carry two at a time. You need an original population to carry all the alleles.
I said it first, see Message 422.
--Percy

This message is a reply to:
 Message 479 by RAZD, posted 04-21-2010 7:49 AM RAZD has seen this message but not replied

Replies to this message:
 Message 483 by PaulK, posted 04-21-2010 9:17 AM Percy has replied

  
PaulK
Member
Posts: 17827
Joined: 01-10-2003
Member Rating: 2.3


Message 483 of 851 (556823)
04-21-2010 9:17 AM
Reply to: Message 482 by Percy
04-21-2010 9:03 AM


Re: Mutations Revisited 2
In fact I said it in Message 46 and Message 111. But In this case I was raising a specific point about interfertility. If it is down to incompatible sets of alleles, and if we accept the standard YEC view that a pairs of an original species on the ark could give rise to a whole taxonomic family we do have to wonder how this could be plausible without seriously compromising the fertility of the original pair or their early offspring. Remember at the start, no alleles can be rare, since each must be represented at least once in half the population !

This message is a reply to:
 Message 482 by Percy, posted 04-21-2010 9:03 AM Percy has replied

Replies to this message:
 Message 485 by Percy, posted 04-21-2010 9:56 AM PaulK has not replied
 Message 498 by Faith, posted 04-21-2010 5:16 PM PaulK has replied

  
ZenMonkey
Member (Idle past 4537 days)
Posts: 428
From: Portland, OR USA
Joined: 09-25-2009


Message 484 of 851 (556834)
04-21-2010 9:48 AM
Reply to: Message 481 by Percy
04-21-2010 8:59 AM


Re: Mutations Revisited 2
Percy writes:
How many closely related species are different in only their allele frequencies? Squirrels? Finches? Cats? Any species at all?
I'd be highly interested in this as well. The ability of speciation arising simply out of allele differences is a part of the train of thought I'm following on the other thread. Probably not the most important part, but one I'd like to be clear on.
And if anyone sees me making any egregious errors in my train of thought over there, please let me know.
Edited by ZenMonkey, : Clarity.

I have no time for lies and fantasy, and neither should you. Enjoy or die.
-John Lydon
What's the difference between a conspiracy theorist and a new puppy? The puppy eventually grows up and quits whining.
-Steven Dutch

This message is a reply to:
 Message 481 by Percy, posted 04-21-2010 8:59 AM Percy has seen this message but not replied

  
Percy
Member
Posts: 22492
From: New Hampshire
Joined: 12-23-2000
Member Rating: 4.9


Message 485 of 851 (556840)
04-21-2010 9:56 AM
Reply to: Message 483 by PaulK
04-21-2010 9:17 AM


Re: Mutations Revisited 2
Okay, I concede, you have priority.
--Percy

This message is a reply to:
 Message 483 by PaulK, posted 04-21-2010 9:17 AM PaulK has not replied

  
misha
Member (Idle past 4654 days)
Posts: 69
From: Atlanta
Joined: 02-04-2010


Message 486 of 851 (556844)
04-21-2010 10:22 AM
Reply to: Message 472 by Peepul
04-21-2010 5:21 AM


Re: Mutations Revisited 2
Peepul writes:
I've been thinking about this in the context of dogs. I suspect (though I don't know) that the massive size variation in dogs is brought about by assortment of alleles rather than mutations. The biggest dogs and the smallest dogs are reproductively isolated from each other - they cannot mate and the small dogs could not bring a mixed puppy to term. Something similar in the wild would generate multiple species.
I see what you're saying. But I think you're wrong in assuming that large dogs and small dogs can't mate. Yes, it is a physical barrier for a large male dog to impregnate a small female dog. And it is a physical barrier for a small female dog to carry a large dog offspring to term. But if you flip the genders reproduction is easily possible. Other than the fact that the large female dog would have to be laying down, is there any reason why a small male dog physically could not impregnate her? Is there any reason why the large female could not carry a small offspring to term?
My wife's old roommate had a rotweiller/dacshund mix. Rotweiller mother, dacshund father. The smaller father had minimal issues impregnating the larger mother and the larger mother had minimal issues carrying the pups to term. The dog grew to be larger than a dacshund but smaller than a rotweiller, a hybrid. A male dog from the litter could easily impregnate another female rotweiller or a female dachshund and then you would have fourth and fifth size variants.
Unless there is a mechanism that would create barriers for a large mother/small father combination then I don't think size could be the lone cause of a speciation event.

This message is a reply to:
 Message 472 by Peepul, posted 04-21-2010 5:21 AM Peepul has replied

Replies to this message:
 Message 517 by Peepul, posted 04-22-2010 3:54 AM misha has replied

  
Peepul
Member (Idle past 5044 days)
Posts: 206
Joined: 03-13-2009


Message 487 of 851 (556852)
04-21-2010 11:46 AM
Reply to: Message 476 by Percy
04-21-2010 5:50 AM


Re: Mutations Revisited 2
quote:
This is the definition Faith is using
Sorry, I hadn't paid enough attention. At least she's made the argument easier!

This message is a reply to:
 Message 476 by Percy, posted 04-21-2010 5:50 AM Percy has replied

Replies to this message:
 Message 488 by Percy, posted 04-21-2010 3:36 PM Peepul has not replied

  
Percy
Member
Posts: 22492
From: New Hampshire
Joined: 12-23-2000
Member Rating: 4.9


Message 488 of 851 (556901)
04-21-2010 3:36 PM
Reply to: Message 487 by Peepul
04-21-2010 11:46 AM


Re: Mutations Revisited 2
Yeah, but WK didn't agree with me. I'm not persuaded that he's approaching it the right way by addressing her paradigm from within, but WK's views have to be given serious consideration.
--Percy

This message is a reply to:
 Message 487 by Peepul, posted 04-21-2010 11:46 AM Peepul has not replied

  
Faith 
Suspended Member (Idle past 1471 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 489 of 851 (556905)
04-21-2010 4:23 PM
Reply to: Message 471 by Wounded King
04-21-2010 4:38 AM


Yes, change, even speciation, without mutations
Have to say I disagree with you here Percy. Within a normal breeding population you are likely to have a spectrum of genetic reproductive compatibility. Given such a situation I can quite readily see a situation where you could produce two daughter populations with distinct strict subsets of the alleles from the parent population but which are not genetically compatible. The loss of alleles or genotypes from the parent population that allowed gene flow between these sub-populations could effectively ensure their reproductive isolation.
Suppose there is a gene Fertile with alleles Fertilea and FertileA in the original population. Homozygotes of either allele can breed with each other and with heterozygotes but neither can produce viable fertile offspring with each other. If this population is split and due to drift the Fertilea and FertileA alleles are fixed in respective populations then we will have established 2 genetically incompatible populations without any further mutations, only through loss of one allele from each population. If we reintroduce the 2 new populations to each other they would not be able to breed.
Off the top of my head I don't know of any examples such as this, but I don't see why it couldn't happen, or how we would not have to consider the resulting sub-populations distinct species.
Given a wide enough panel of genes with alleles giving rise to hybrid inviability we might even see a situation like the Greenish Warbler ring species arising through such a reductive form of speciation.
Thank you for the acknowledgment of a possibility that fits with what I'm saying and a good example.

This message is a reply to:
 Message 471 by Wounded King, posted 04-21-2010 4:38 AM Wounded King has not replied

  
Faith 
Suspended Member (Idle past 1471 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 490 of 851 (556906)
04-21-2010 4:26 PM
Reply to: Message 472 by Peepul
04-21-2010 5:21 AM


reproductive isolation by whatever means
physical and behavioural differences that prevent reproduction of one group with another are sufficient to make them into different species, aren't they?
That's how I've always understood it. I don't insist on HOW reproductive isolation occurs, I'm trying to use the standard definitions of these things and this is one way species are considered to form.

This message is a reply to:
 Message 472 by Peepul, posted 04-21-2010 5:21 AM Peepul has not replied

  
Faith 
Suspended Member (Idle past 1471 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 491 of 851 (556907)
04-21-2010 4:30 PM
Reply to: Message 471 by Wounded King
04-21-2010 4:38 AM


Re: Mutations Revisited 2
I think RAZD's distinction was that you can't get a genetically reproductively isolated population if another population still exists which retains all the parent populations allelic diversity. In other words, if either of my two Fertilea/a and FertileA/A sub populations were reintroduced to the parent population which still had FertileA/a individuals and those of their own genotype in it then they could introgress.
I'm aware of this possibility which is why I try to remember to always specify that reproductive isolation is necessary for what I'm saying to occur. This is why I try to always say "barring gene flow" and that sort of thing. I'm trying to focus on what happens with isolated populations inbreeding on their own after a migration or series of migrations from the original population.

This message is a reply to:
 Message 471 by Wounded King, posted 04-21-2010 4:38 AM Wounded King has not replied

  
Faith 
Suspended Member (Idle past 1471 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 492 of 851 (556908)
04-21-2010 4:32 PM
Reply to: Message 473 by Percy
04-21-2010 5:26 AM


Re: Mutations Revisited 2
She doesn't understand much of what reads both here and elsewhere. A great recent example is when she stated that she thought we'd been saying that mutations don't change the basic genomic structure of species. How basic a misunderstanding is that!
There were two or three such statements and I hope I can eventually find them to see if what you are saying applies or not, something to the effect that of course mutations are going to be compatible with the species, and I don't remember why it was said, apparently in answer to something I said.

This message is a reply to:
 Message 473 by Percy, posted 04-21-2010 5:26 AM Percy has replied

Replies to this message:
 Message 494 by Percy, posted 04-21-2010 4:52 PM Faith has not replied
 Message 507 by RAZD, posted 04-21-2010 8:10 PM Faith has not replied

  
Faith 
Suspended Member (Idle past 1471 days)
Posts: 35298
From: Nevada, USA
Joined: 10-06-2001


Message 493 of 851 (556909)
04-21-2010 4:51 PM
Reply to: Message 476 by Percy
04-21-2010 5:50 AM


allele subsets? No, new allele frequencies.
It would be better to refute Faith's argument using the 'interbreeding populations' definition of species. Although there is no formal agreement to it, that's the most commonly used definition.
This is the definition Faith is using, and if you don't peer under the hood at the genetic underpinning of the phenotypes then you'd might conclude she's right. But closely related populations differ not only in the alleles for each gene, but also in terms of which genes they have and even which chromosomes they have.
This sounds to me perfectly compatible with my model and in fact good evidence for it. Such differences certainly suggest problems with interbreeding and also suggest that reduced genetic diversity plays a part in such a situation.
I think examples of closely related populations where the differences are in allele subsets only are rare. Even WK couldn't think of any examples.
WK very nicely showed how interbreeding problems could occur simply from allele loss without mutations and lead to speciation.
I'm always saying that the smaller the daughter population the more apparent should be the effects, certainly the smaller the genetic diversity in the new population, certainly the emergence of a distinct new overall look from the new combination of a reduced number of traits available. I've also referred to the fairly equal splitting of an initial population as a probably less dramatic way the same events come about -- which is why I like the smaller-population example to make the point clearer.
And I've also never said I expect speciation with the first or second migration though it COULD happen that interbreeding on a genetic basis could stop -- I'm including ALL the ways you can get reproductive isolation such as geographic barriers and behavioral barriers AND genetic barriers, because my point is all about what happens when you HAVE reproductive isolation, not how you get there.
And again, I also am not making as big a point about speciation itself as some of you are. I started emphasizing it when people complained that where I was talking about evolution I seemed to mean speciation and I could see their point, although I also want to make the point that macroevolution depends on speciation, as some have acknowledged. I see speciation as the necessary end result of these processes if you're going to have macroevolution, but it's the trend up to and including that point that I'm trying to stay focused on -- the reduction in genetic diversity that allows the emergence of new phenotypes which become the basis of new varieties.

This message is a reply to:
 Message 476 by Percy, posted 04-21-2010 5:50 AM Percy has replied

Replies to this message:
 Message 495 by Percy, posted 04-21-2010 5:04 PM Faith has not replied

  
Percy
Member
Posts: 22492
From: New Hampshire
Joined: 12-23-2000
Member Rating: 4.9


Message 494 of 851 (556910)
04-21-2010 4:52 PM
Reply to: Message 492 by Faith
04-21-2010 4:32 PM


Re: Mutations Revisited 2
Faith writes:
There were two or three such statements and I hope I can eventually find them to see if what you are saying applies or not, something to the effect that of course mutations are going to be compatible with the species, and I don't remember why it was said, apparently in answer to something I said.
I wouldn't waste time and effort reconciling your different understandings at different times. Just keep working on improving your understanding, which is what all of us are doing here. Anyone who's understandings of everything are the same as even just a year ago isn't learning anything.
Anyway, I commented on this back in Message 456:
Percy in Message 456 writes:
Faith writes:
It's been emphasized by a few here that mutations always accord with the character of the species anyway...
No one on this thread has even remotely suggested this. You're probably thinking of explanations about beneficial mutations tending to be in very tiny undetectable steps. Instances of easily apparent beneficial mutations should be rare.
Reproduction is almost invariably imperfect. There's not really any such thing as "the character of the species." All you can talk about is the current moment in a process of continuous accumulating change over time. Species classifications are just human beings imposing their own classification systems at a static moment in time.
--Percy

This message is a reply to:
 Message 492 by Faith, posted 04-21-2010 4:32 PM Faith has not replied

  
Percy
Member
Posts: 22492
From: New Hampshire
Joined: 12-23-2000
Member Rating: 4.9


(1)
Message 495 of 851 (556911)
04-21-2010 5:04 PM
Reply to: Message 493 by Faith
04-21-2010 4:51 PM


Re: allele subsets? No, new allele frequencies.
Faith writes:
It would be better to refute Faith's argument using the 'interbreeding populations' definition of species. Although there is no formal agreement to it, that's the most commonly used definition.
This is the definition Faith is using, and if you don't peer under the hood at the genetic underpinning of the phenotypes then you'd might conclude she's right. But closely related populations differ not only in the alleles for each gene, but also in terms of which genes they have and even which chromosomes they have.
This sounds to me perfectly compatible with my model and in fact good evidence for it. Such differences certainly suggest problems with interbreeding and also suggest that reduced genetic diversity plays a part in such a situation.
I'm sure all of us just collectively threw up our hands in frustration.
Faith, your position is that species are created through allele reduction. You deny mutation any significant role. Without mutations you cannot get differences in which genes and chromosomes each population has. In your model the genes and chromosomes between the two populations will remain forever the same.
Mutations are the only way that changes in the character and number of genes and chromosomes can come about. You can't say that your model is "perfectly compatible" if it denies mutations any significant role. Without mutations your model isn't even a teensy tiny bit compatible.
--Percy

This message is a reply to:
 Message 493 by Faith, posted 04-21-2010 4:51 PM Faith has not replied

  
Newer Topic | Older Topic
Jump to:


Copyright 2001-2023 by EvC Forum, All Rights Reserved

™ Version 4.2
Innovative software from Qwixotic © 2024