Natural Selection - not natural?

I definitely need to reexamine the study in detail. I had actually forgotten about the delayed inheritance for some reason and was thinking in these terms when I made the single generation claim - definitely my mistake but perhaps explains my reasoning. Right - I was writing about the hypothetical Mendelian inheritance (vs. the actual study), so I don't think we have an argument on this point...

I don't think there is really anything non-Mendelian about it simply because it is a maternal effect gene.It is a really interesting paper, and gives some really good evidence for one of the species having 'speciated' in as much as becoming reproductively isolated from small peripheral populations and the late reabsorbing those populations and becoming in effect a true species with a polyphyletic origin rather than just a polyphylatic species due to the different species being cryptic.This only applies in the loose sense of speciation of course as there doesn't appear to have been any post-mating or physiological block to reproduction.TTFN,WK