The first upright ape was also human. In the millions of years that followed, new species branched off and abandoned their upright posture to descend to what we now call "ape."
Nonsense. The first upright ape, according to this find was Morotopithecus and its close cousins had the potential ability of bipedal locomotion. It is possible that the other great apes had an ancestor that was able to walk upright. I have no reason to suspect that this fact, should it be true, means that current Evolutionary Theory as it now stands cannot explain it.
How could all apes bar one end up with the hunched over character
That question can be asked without this being an issue, we are uniquely bipedal -why? The question is why did evolution modify this trait in us so that we are bipedal specialists (Morotopithecus would probably have not been exclusively bipedal from what I have read), when it modified all the other surviving lineages to have less bipedalism?
how could our spine be so screwy if it had evolved for that long
Our spines of course, have been evolving for much longer. However, there is the possibility that our lineage flirted with both bipedalism and quadrapedalim for some time before evolving into the specialists we are.
Edited by Modulous, : No reason given.