Dimensions | | No. of local populations |

| *n*_{X} | No. of individuals in population *X* |

| | No. of traits |

| *n*_{ℒ} | No. of loci |

Coancestry coefficients | Matrix **θ**, with elements θ_{ii}_{′} | Coancestry coefficient = probability that randomly chosen alleles for individuals *i* and *i*′ are identical by descent for a neutral locus |

| Matrix , with elements | Population-to-population coancestry |

| | Average population-to-population coancestry |

| | Within-population coancestry |

| | Average within-population coancestry |

| | Average self-coancestry in population *X* |

| | Average self-coancestry |

Distribution of alleles | *x*_{ijku} | Indicator function for the allele *k* in locus *j* of individual *i* being of the allelic type *u* |

| *p*_{ju} | The frequency of allele *u* in locus *j* in the ancestral population |

Genetic values | *v*_{jum} | Additive value of allele *u* in locus *j* for trait *m* |

| | Additive value of trait *m* for individual *i* |

| **a**_{i} | Vector of additive values (of all traits) for individual *i* |

| | Mean additive value in population *X* |

| | Mean additive value |

Genetic variance–covariance matrices | | Matrix of genetic variances and covariances in the ancestral population |

| G_{X} | Matrix of genetic variances and covariances in population *X* |

| | Matrix of genetic variances and covariances among populations |