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Author Topic:   Question on evolutionary Rates
Chiroptera
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Message 16 of 47 (393766)
04-06-2007 9:20 PM
Reply to: Message 15 by ICANT
04-06-2007 9:17 PM


Re: Re-Every 200,000 years
quote:
But if you say OK forget all those problems we had a new species every 200,000 years you would have the 7500 levels.
Even assuming that the 200,000 year per speciation is correct (and I doubt that it is), then what is the problem with 7500 "levels"?

Actually, if their god makes better pancakes, I'm totally switching sides. -- Charley the Australopithecine

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RAZD
Member (Idle past 1432 days)
Posts: 20714
From: the other end of the sidewalk
Joined: 03-14-2004


Message 17 of 47 (393772)
04-06-2007 9:59 PM
Reply to: Message 14 by ICANT
04-06-2007 9:13 PM


running changes
If they could have provided a detailed trail with the formas forking off and becoming some other family of creature they would have been heros.
Why? There is plenty of evidence for this happening, so it would not add anything new to the concept of evolution or the evidence for it.
The foraminifera are also an order not just a species, so there are changes at the family level too (between species and order).
If you want an example of species evolving into a substantially different species try horses. Part of this depends on what you think is distinctive enough of a difference ...
abe: this is copied from another thread that is not open to general discussion.
No webpage found at provided URL: distinctively - adv.
in an identifiably distinctive manner; "the distinctively conservative district of the county"
No webpage found at provided URL: distinctly - adv.
1: clear to the mind; with distinct mental discernment; "it's distinctly possible"; "I could clearly see myself in his situation" [syn: clearly] 2: in a distinct and distinguishable manner; "the subtleties of this distinctly British occasion" 3: to a distinct degree; "urbanization in Spain is distinctly correlated with a fall in reproductive rate"
No webpage found at provided URL: dis·tinc·tive - adj.
1. Serving to identify; distinguishing: distinctive tribal tattoos. See Usage Note at distinct.
2. Characteristic or typical: “Jerusalem has a distinctive Middle East flavor” (Curtis Wilkie).
3. Linguistics. Phonemically relevant and capable of conveying a difference in meaning, as nasalization in the initial sound of mat versus bat.
We can talk about horses and the distinctive development of the modern horse and single-toe hoof from the splayed toed dog sized "eohippus":
http://www.flmnh.ufl.edu/...ci/vertpaleo/fhc/relatives10.htm (2)
quote:
Look at how the bones in horse feet have changed over time.
They became longer and more streamlined, enabling horses to run faster to avoid predators.
Check the above link to see images of the legs of four different horse ancestors. You can see a splayed toe stance for Hyracotherium and Miohippus but a single toe stance in Merychippus and Equus.
http://www.geocities.com/...rk/7841/horse_evol/eohippus.html (1)
quote:
Eohippus was a descendent of the Condylarth, a dog-sized, five-toed creature that lived about 75 million years ago. It lived during the early Eocene period, which took place 50 to 60 million years ago. Eohippus, which means "dawn horse," stood about twelve to fourteen inches at the shoulder and weighed about twelve pounds. It looked nothing like a horse. It had an arched back, short neck, short snout, short legs, and a long tail. Its color probably most resembled that of a deer, a darker background with lighter spots.
The legs of Eohippus were flexible and rotating with all major bones present and unfused. It had a choppy, up-down gait and was not very fast. There were four toes on each front foot and three toes on the hind. The vestigial toes - two on the front feet and one on the hind - were still present.
It had a small brain and low-crowned teeth with three incisors, one canine, four distinct premolars, and three "grinding" molars in each side of each jaw. Browsing on fruit and fairly soft foliage, Eohippus probably lived in an environment with soft soil, the kind found on jungle floors and around the edges of pools. Since Eohippus walked on the pads of its feet, it was able to cross wet, marshy ground without much difficulty.
The coloration is pure speculation, of course, but the size and stance are based on the physiology of the skeleton. Now lets also look at the Condylarth:
Paleocene mammals of the world (3)
quote:
Back in the northern hemisphere, another family of condylarths, the Phenacodontidae, may include the ancestors of a more familiar ungulate order: The odd-toed ungulates or Perissodactyla, represented by horses, rhinos and tapirs in the recent fauna. Historically, phenacodontids form the core of the Condylarthra. Well-preserved skeletons are known for the type genus Phenacodus, which is a good model of an ancestral ungulate with beginning adaptations for running. Unlike arctocyonids, periptychids or mioclaenids, the phenacodontids are not part of the first wave of condylarths that populated North America. They first appear with the fox-sized Tetraclaenodon in the middle Paleocene of that continent. The appearance of the more advanced phenacodontids Phenacodus and Ectocion marks the beginning of late Paleocene time in North America. The type genus Phenacodus covers the large size range of phenacodontids and includes roughly sheep-sized animals. Members of the genus Ectocion were usually smaller, with a body mass of only 3 kg in the smallest species, but there is some overlap in size between the two genera. Phenacodontids were the dominant mammals in the latest Paleocene of North America and account for up to 50% of all mammal specimens in faunas of that age.

Figure 5: Reconstruction of the late Paleocene to middle Eocene Phenacodus, a sheep-sized herbivore with improved capabilities for running. From Savage & Long (1986).
This is the most "horse-like" image from this site, and it looks much more like a dog than a horse eh? Of course the coloration and fur are speculation, but the size and stance are again based on the physiology of the skeleton.
To see what the skeletons looked like for the four species used for the leg examples at the start we go to:
http://www.flmnh.ufl.edu/natsci/vertpaleo/fhc/Stratmap1.htm (2)
quote:
(NOTE: LINKS INTENTIONALLY BROKEN - this image is copied from the original to save bandwidth, go to the original Stratmap for the links to work).
(Hyracotherium)- This small dog-sized animal represents the oldest known horse. It had a primitive short face, with eye sockets in the middle and a short diastema (the space between the front teeth and the cheek teeth).
Although it has low-crowned teeth, we see the beginnings of the characteristic horse-like ridges on the molars.
Hyracotherium is better known as "eohippus" - which means "the dawn horse." The name also refers to the fact that it lived during the Eocene.
(Miohippus) - Species of Miohippus gave rise to the first burst of diversity in the horse family. Until Miohippus, there were few side branches, but the descendants of Miohippus were numerous and distinct. During the Miocene, over a dozen genera existed.
Fossils of Miohippus are found at many Oligocene localities in the Great Plains, the western US and a few places in Florida. Species in this genus lived from about 32-25 million years ago.
(Merychippus) - Merychippus represents a milestone in the evolution of horses. Though it retained the primitive character of 3 toes, it looked like a modern horse. Merychippus had a long face. Its long legs allowed it to escape from predators and migrate long distances to feed. It had high-crowned cheek teeth, making it the first known grazing horse and the ancestor of all later horse lineages.
Fossils of Merychippus are found at many late Miocene localities throughout the United States. Species in this genus lived from 17 - 11 million years ago
(Equus) - Equus is the only surviving genus in the once diverse family of horses. Domesticated about 3,000 years ago, the horse had a profound impact on human history in areas such as migration, farming, warfare, sport, communication and travel.
Species of Equus lived from 5 million years ago until the present. Living species include horses, asses, and zebras. Fossils of Equus are found on every continent except Australia and Antarctica.
I selected the same species as were listed for the legs above for convenience here - on the original link you select by clicking on the skulls.
So we have a sequence of species that starts with one standing on the fleshy pads of several splayed toes to the modern species that stands not just on one toe but on the toe-nail of that single toe. But that is not all:
http://muextension.missouri.edu/...agguides/ansci/g02740.htm (4)
quote:
A horse's hoof is composed of the wall, sole and frog. The wall is simply that part of the hoof that is visible when the horse is standing. It covers the front and sides of the third phalanx, or coffin bone. The wall is made up of the toe (front), quarters (sides) and heel.
The wall of the hoof is composed of a horny material that is produced continuously and must be worn off or trimmed off. The hoof wall does not contain blood vessels or nerves. In the front feet, the wall is thickest at the toe; in the hind feet the hoof wall is of a more uniform thickness. The wall, bars and frog are the weight-bearing structures of the foot. Normally the sole does not contact the ground.
As weight is placed on the hoof, pressure is transmitted through the phalanges to the wall and onto the digital cushion and frog. The frog, a highly elastic wedge-shaped mass, normally makes contact with the ground first. The frog presses up on the digital cushion, which flattens and is forced outward against the lateral cartilages. The frog also is flattened and tends to push the bars of the wall apart (Figure 3). When the foot is lifted, the frog and other flexible structures of the foot return to their original position.
When the foot is placed on the ground, blood is forced from the foot to the leg by the increase in pressure and by the change in shape of the digital cushion and the frog. The pressure and the change in shape compress the veins in the foot. When the foot is lifted, the compression is relieved and blood flows into the veins again. In this way, the movement of these structures in the hoof acts as a pump.
This is much more difference in a feature than "just an increase in length" (as in an elephants trunk), it is a totally different structure to stand on (eohippus stood on his toes pads, equus stands on a hoof which not only is not a toe pad, but a feature that wasn't present in the eohippus) and it incorporates a new {added\changed} structure to increase blood flow by acting as a secondary pump.
Not only that the effect of changing the foot structure from a flat footed splayed toed eohippus to the single toed equus also involves standing the foot up on the tip of the toe and using each of the bones between the tip and the heel to effectively make the leg longer for faster running while also making it more flexible than just adding length to the bones of the leg. Probably useful for getting through tight spots and to keep from tripping ... it certainly helps horses jumping in shows from hitting that top bar.
Totally different foot structure, coupled with totally different leg structure (with some ex toe bones now effectively used as leg bones).
The question again is how much change is enough? Try walking around the house on the tip of one toe, then compare your foot to that of eohippus.
Enjoy.


References:
  1. Anonymous, "Evolution of the Horse: Eohippus" Sarah's Horse Farm, Updated 10 Aug 2000, Accessed 12 Jan, 2007
  2. Anonymous, "Fossil Horse Cybermuseum" The Florida Museum of Natural History, Updated 8 Jan 2007, Accessed 12 Jan, 2007
  3. Jehle, Martin, "Condylarths: Archaic hoofed mammals" Paleocene mammals of the world, Updated ?, Accessed 12 Jan, 2007
  4. McClure, Robert C. et al "Functional Anatomy of the Horse Foot" Department of Veterinary Anatomy, College of Veterinary Medicine, Updated 10 May 2006, Accessed 12 Jan, 2007
Edited by RAZD, : abe

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crashfrog
Member (Idle past 1494 days)
Posts: 19762
From: Silver Spring, MD
Joined: 03-20-2003


Message 18 of 47 (393785)
04-07-2007 12:20 AM
Reply to: Message 15 by ICANT
04-06-2007 9:17 PM


Re: Re-Every 200,000 years
What I am saying at the 200,000 year rate you could only have 7500 levels.
Sure. At the bottom level, though, you'd have 2^7500 different species.
It's all but impossible to imagine how many species that is. That's more species than there are atoms in the entire visible universe. That's more than the number of atoms in 100 visible universes exactly like ours.
Is it really so unreasonable to believe that Homo sapiens would be just one of those species? I don't see that it is. It's more unreasonable to believe that Homo sapiens wouldn't be one of those species.
Then when they started to mate you have the added problem of mating.
Sexuality, as near as we can tell, emerged quite early in evolutionary history, so I don't see how it's a problem. We have so many sexual organisms because they're all the decendants of the first sexual species, not because they all evolved sexuality independantly. (It's possible, now that I think about it, that sexuality may have evolved two or three times independantly, since there's at least two or three different chromosomal schemes for sex determination that we know about.)
That problem had to be solved along the way somewhere.
It's not a hard problem to solve. Consider a genetic factor that's passed on only 50% of the time. If you have it, you're Sex A. If you lack it, you're Sex B.
Of course, that's not the only way to do it. You can have more than 2 genders, and many species do. But specialized sex structures and body types come long after the genetic basis for sex in evolutionary history. It's interesting but it's not a major problem.

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jar
Member (Idle past 421 days)
Posts: 34026
From: Texas!!
Joined: 04-20-2004


Message 19 of 47 (393786)
04-07-2007 12:24 AM
Reply to: Message 18 by crashfrog
04-07-2007 12:20 AM


Re: Re-Every 200,000 years
Of course, that's not the only way to do it. You can have more than 2 genders, and many species do. But specialized sex structures and body types come long after the genetic basis for sex in evolutionary history. It's interesting but it's not a major problem.
And you have species where sex is determined by external conditions and species where sex changes with maturity and species that change sex and species were a single organism is both sexes and ...

Aslan is not a Tame Lion

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Wounded King
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Posts: 4149
From: Cincinnati, Ohio, USA
Joined: 04-09-2003


Message 20 of 47 (393819)
04-07-2007 1:01 PM
Reply to: Message 10 by RAZD
04-06-2007 2:31 PM


Foraminifers are sexual and asexual, but not conscious (so sexual selection not a factor).
This a contentious issue, some people argue that plants can have sexual selection, so I can't imagine they would except Foraminifers.
TTFN,
WK

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ICANT
Member
Posts: 6769
From: SSC
Joined: 03-12-2007
Member Rating: 1.6


Message 21 of 47 (393852)
04-07-2007 9:33 PM
Reply to: Message 18 by crashfrog
04-07-2007 12:20 AM


Re: Re-Every 200,000 years
It's all but impossible to imagine how many species that is.
Number reaches Infinity.

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Dr Adequate
Member (Idle past 312 days)
Posts: 16113
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Message 22 of 47 (393853)
04-07-2007 9:38 PM
Reply to: Message 21 by ICANT
04-07-2007 9:33 PM


Re: Re-Every 200,000 years
A while ago you didn't believe that speciation didn't happen at all. Now you think that the number of species should "reach infinity".
Here's a tip: try relating your statements to reality.

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ICANT
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Posts: 6769
From: SSC
Joined: 03-12-2007
Member Rating: 1.6


Message 23 of 47 (393858)
04-07-2007 11:22 PM
Reply to: Message 22 by Dr Adequate
04-07-2007 9:38 PM


Re: Re-Every 200,000 years
Here's a tip: try relating your statements to reality.
Do the math yourself start with 2 double for 7500 times.
A while ago you didn't believe that speciation didn't happen at all.
????

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NosyNed
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Posts: 9004
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Message 24 of 47 (393867)
04-08-2007 3:25 AM
Reply to: Message 23 by ICANT
04-07-2007 11:22 PM


Rice grains
Do the math yourself start with 2 double for 7500 times.
And if I put a grain of rice on the first square of a chess board, two on the next; 4 on the third ....
but what does that have to do with reality?

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Zucadragon
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Posts: 69
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Message 25 of 47 (393868)
04-08-2007 3:33 AM


ICANT its not so much a number issue really, what you're doing now is saying that the change from the first eukaryotic organismen to a human is a more complicated one then the change from a prokaryotic organism to an eukaryotic one.. Or atleast on the same level.
But that original statement is wrong in a very important way. And I'll provide a little example to explain it.
The Drosophilia fly is a bug of which we know all genetic information, we also know that (and this is well known for anyone dealing with the fly, ask around biologists and they'll explain it to you) the fly has many genes that are closely related to human genes, around 78% I hear and read in a lot of places. Yet if you look at it, it still looks like a simple bug.
And thats where the point lies, you can think about the first multicellular organismes and say "those would be so simple". But those first multicellular organismes would already have many of the genes that the humans would have in the future as well, minus and plus extra changes.. Just like the fly now, would those first organismes have many of the basic genes..
Which would take it less time in "speciation events" to get to the stage of what we call a human now.
Does that make sense ?
Add to this one more bit of information.. The genes from a prokaryote to an eukaryote was a big change, it wasn't something present in the organism and thus had to be created during a long process.. When the genes were there, you'd have eukaryotes, it is ofcourse a smaller step to go from one eukaryote to a different one..

  
RAZD
Member (Idle past 1432 days)
Posts: 20714
From: the other end of the sidewalk
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Message 26 of 47 (393966)
04-08-2007 7:33 PM
Reply to: Message 20 by Wounded King
04-07-2007 1:01 PM


This a contentious issue, some people argue that plants can have sexual selection, so I can't imagine they would except Foraminifers.
True. And one could argue that sexual selection also goes on in mosquitoes and the like via variations in the sexual organs\fit.
Perhaps what I mean is conscious\unconscious selection - based on some mental perception.
So the question is where does run-away sexual selection fit on the conscious\unconscious scheme of things? This would represent the fast-tracked selection process that, imh(ysa)o, would show a faster than normal rate of fixing selected mutations.
Thanks.

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ICANT
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Posts: 6769
From: SSC
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Message 27 of 47 (393990)
04-08-2007 10:57 PM
Reply to: Message 26 by RAZD
04-08-2007 7:33 PM


Re-Numbers
Hi RAZD in your earlier post you gave a chart on the horse.
It shows 15 different species within 8 levels. The time frame for this was 55 million years. Now this was after science says we have a horse. The average speciation rate per level would be 1 every 550,000 years. Almost 3 times as long as the Foraminifers.
Merychippus 17 - 11 million years ago
Pliohippus 12-6 million years ago.
Update: Over the years, refinements in our understanding of the Pliocene have resulted in the shrinking of that epoch. It is now believed to have covered a period ranging from about 5.3 to 1.75 million years ago.
Dinohippus appears to be the closest relative to Equus. It seems to be an intermediate between Pliohippus and Equus. In fact, the genus was established from species that were initially included in Pliohippus (e.g. P. leidyanus).
I have a problem understanding how Dinohippus can be an intermediate between Pliohippus and Equus when Pliohippus didn't exist until 5.3 million years ago. If he is then we have an unaccounted for time period of 5.7 million years.
would show a faster than normal rate of fixing selected mutations.
It does not appear that the horse went very fast.
The question remains with only 7500 levels how did modern man evolve from the Eukaryotic Cells.
If he did, how did he do it without leaving a lot of fossils?

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fallacycop
Member (Idle past 5548 days)
Posts: 692
From: Fortaleza-CE Brazil
Joined: 02-18-2006


Message 28 of 47 (394002)
04-09-2007 1:04 AM
Reply to: Message 27 by ICANT
04-08-2007 10:57 PM


I have a problem understanding how Dinohippus can be an intermediate between Pliohippus and Equus when Pliohippus didn't exist until 5.3 million years ago. If he is then we have an unaccounted for time period of 5.7 million years.
How is the 5.3 million date a problem? Intermediate simply means in between.
The question remains with only 7500 levels how did modern man evolve from the Eukaryotic Cells.
I don`t think this counting of levels make any sense, but even assuming the method has something to it, how isn`t 7500 levels enought?

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RAZD
Member (Idle past 1432 days)
Posts: 20714
From: the other end of the sidewalk
Joined: 03-14-2004


Message 29 of 47 (394016)
04-09-2007 8:16 AM
Reply to: Message 27 by ICANT
04-08-2007 10:57 PM


apples and oranges
It shows 15 different species within 8 levels.
It shows 15 different genera (several species). That's why the different first part of the scientific names. Look at Equus:
http://www.flmnh.ufl.edu/natsci/vertpaleo/fhc/equus1.htm
quote:
Equus is the only surviving genus in the once diverse family of horses.
Living species include horses, asses, and zebras. Fossils of Equus are found on every continent except Australia and Antarctica.
And there are extinct species within Equus. One within modern times (recorded history). Not all genera represented by multiple species (eg - man) but most are (else why create a different genus? It's an arbitrary distinction).
It does not appear that the horse went very fast.
When you include the species within the genus categories then it does appear that horse was faster than shown here, though the chart isn't detailed to the species level (the way foraminifera were, nor is the foraminifera data broken down by genera).
It does not appear that horse ancestors engaged in the type of run-away sexual selection for specific traits that I was talking about either -- there is no sexual dimorphism beyond a slight difference in size (and that is due to dominant male herding behavior) -- and no "peacock tail" features. When we look at man we do see sexual dimorphism and "peacock tail" features: bare-appearing skin, long head hair, large creative brain, large sexual organs, and the like -- more extreme than in any other ape or primate, and some (bare skin, long hair, large brain) extreme to the point of threatening survival of those with the features. This to me is a significant element of human evolution since the common ancestor with chimps.
The question remains with only 7500 levels how did modern man evolve from the Eukaryotic Cells.
The Abbott and Costello (and 3 stooges, and other vaudeville acts) way
Step by step, slowly he turns ....
Enjoy.
Edited by RAZD, : fix quote and plural

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Allopatrik
Member (Idle past 6214 days)
Posts: 59
Joined: 02-07-2007


Message 30 of 47 (394017)
04-09-2007 9:15 AM
Reply to: Message 29 by RAZD
04-09-2007 8:16 AM


Plural of Genus
Just a gentle point: the plural of genus is genera.
A

Natural Selection is not Evolution-- R.A. Fisher

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