Page's misuse of Haldane's Dilemma

Interesting stuff. Largely irrelevant, I suspect, but interesting.I'd be interested in how the graph was arrived at - criteria for defining benefit, how mutations were observed, in what species, that sort of thing.How would it class say a mutation to greater body weight by the increased adiposity in a mammal? Detrimental - making escape from predators more difficult? Beneficial - better able to survive a harsh winter? Or beneficial today but detrimental tomorrow?I'm skeptical about graphing a qualitative judgement.