Quick,
I can come at the bat from Rayn's frog. For HOw does the frog get check mated when the bat can hear the call type and so the female perfers this same sound? whether by echo-location, biaurilization etc or not@
William Provine claims to be a scholar of Sewall Wright yet at one point in Wright's reasoning he asserts that one need think of the a trial and error process for the whole organism, if one is going to apply the shifting balance theory, and I did indeed think of one fro the frog which being preyed on by "your" bat would in co-evolution speak to some of the same variables in any model of the process whether in the trail or error of mankind trying to understand the past that can not be repeated.
Since sound can travel either longitudianlly OR transversely, I reasonsed that the interaction of any sound with the environement can move both longitudinally or trasversely depending on the barrier being percentage wise composed of solids liguids and gases and that so symbolized the bat may have found the "solution" of this engineerable problem frogs in general "solved".
That is the qualititave version without the quality of the bat involved which would be required to complete as well.