mick,
I've read just about all of this thread, and I'm excited because I actually have a question. The questions was already asked by Jianyi in the course of the thread, but I'd like to ask again. Wounded King
may have answered the question in passing, in
message 93, by saying:
Wounded King writes:
The existence of many species showing spectra of interfertility demonstrate that there is a variable range of interfertility. Why do you feel there is some barrier to complete interfertility between 2 populations developing when there are so many populations in which this process is already underway?
But I'm not sure. I had a harder time following that thread of investigation than yours, so I'm addressing this question to you, and not Wounded King. But if WK can clear this up... I'd be delighted to read the response.
First I wanted to say, the definition of "speciation" being used by Jianyi (and by me here) is (I think) an inability to reproduce with organisms of it's parent's species. OK. Here's my question:
What is the mechanism for speciation in neo-darwinian evolution theory?
Your argument, if I understand it correctly, is based on population genetics and collecting mutations based on reproductive isolation. Maybe I'm wrong, but (given this absolute, all-or-nothing view of speciation) I think that doesn't answer the question, and here's why:
Let's say a population of a single species (S1) is split into two groups (G1, G2), i.e. become reproductively isolated. They start as the same species, S1. One of the groups (G2) changes (eventually) to a new species (S2).
I THINK I understand the population genetics view, but here's my question: every organism in G2 is EITHER in S1 or S2, right? If that's right, then a single organism can either reproduce ONLY with S1 or S2. If that's right, then the population genetics mechanism is simply a larger and larger proportion of organisms in G2 being S2, and less being S1.
If that's right, then the question is, how does the S2 population come to be? I think it's clear that the population can increase via NS.
Let's say just one S2 organism is born. It can't reproduce, right? There's no S2 organisms. So, it dies without reproducing. It can't be like that. Then, it must be that TWO S2 organisms were born at the same time in order to reproduce. Well, that's exceedingly unlikely, right?
If all of this is the case, then HOW, at the INDIVIDUAL level, does the S2 population begin? In this way, Jianyi's hypothesis... at least answers the question.
Now, maybe this view on speciation is in error. Maybe an organism can belong to S1 AND S2 AT THE SAME TIME (meaning it can reproduce with BOTH S1 and S2 at the same time, thus creating a "bridge" between the species). In this case, thinking about species as "organisms which cannot sexually reproduce and produce viable offspring" may not be a valid way of thinking. I'm not saying that you suggested it IS a valid way of thinking, just that some people use it as a definition of species / speciation (including, in my own thoughts, me, and also Jianyi).
As you can tell from my writing, and my previous questions that you've graciously answered, I'm still "out of the loop" on this stuff. But I feel there's been a mismatch between Jianyi's statements and yours, and the movites I see behind Jianyi's hypothesis are ones that I don't have an answer to, and that I couldn't derive an answer from your posts from either.
Anyway, thanks for reading, and thanks in advance for any clarification.
Ben