First let us remind ourselves what Richard Dawkins said...Obviously it is Richard Dawkins not me who forgot on replication and heritable traits
You quoted the conclusion not the setup - did you think I wouldn't notice or something. The set up is as follows:
quote:
Suppose it is important in a really successful crew that the rowers should coordinate their activities by means of speech. Suppose further that, in the pool of oarsmen at the coach's disposal, some speak only English and some speak only German. The English are not consistently better or worse rowers than the Germans. But because of the importance of communication, a mixed crew will tend to win fewer races than either a pure English crew or a pure German crew.
The coach does not realize this. All he does is shuffle his men around, giving credit points to individuals in winning boats, marking down individuals in losing boats...What will emerge as the overall best crew will be one of the two stable states--pure English or pure German, but not mixed. Superficially, it looks as though the coach is selecting whole language groups as units. This is not what he is doing. He is selecting individual oarsmen for their apparent ability to win races... Selection at the low level of the single gene can give the impression of selection at some higher level.
So do we have replication? Yes. The 'genes' replicate in that the rowers are moved on into the next generation of boats. They are also shuffled (genetic recombination). The boats inherit the genes from the last race. The coach selects consistently winning individuals to be 'passed on' (he is natural selection in this analogy) and consistently losing individuals to not be passed on. By doing this he can construct a solid team without ever knowing that he is selecting based on language.
In the next generation "individual oarsman" will be in completely different team of oarsmen.
Yes - the shuffling of genes is more extreme than in nature, but the analogy is to show how the genes are being selected for their ability to work well with the other genes in the pool to produce a good team on average.
So each allele will be sitting in the next generation in almost completely different set of alleles (crew). Each individual (boat, crew) is unique genotyp. There are not the same genotypes - except true twins. There are always different phenotypes
At first - but since the coach is selecting his team some alleles will be on less teams and some will be on more teams as time goes on. Gene frequency, in the sense conveyed in the analogy, is shifting.
Because of this the effect of each individual allele will be different in each generation and each genotype. The effect of an allele depends predominantly on the mix of other alleles. The influence of an allele sitting in unique set of other different alleles to the phenotype and its fitness is always different.
Precisely! So an allele needs to be on a consistently winning team. If there are 10 English men and 4 Germans in the pool, the Germans will often be on teams with English men and so will often lose and get marked down, making them less likely to be picked for other teams in the future.
There is no doubt some alleles are detrimental and will be removed by NS. But other alleles and genes influence each other and the outcome depends on their mix which is in each generation different. Obviously there is no Dawkins hypothetical relation between an individual allele and fitness of the phenotype.
Dawkins puts forward the view point that genes that work together well to build better 'boats' or phenotypes will be selected for and increase in frequency.
Edited by Modulous, : No reason given.