I think there is a problem with this. There isn't any, that I know of, good clear definition of trait. It is, AFAIK, used loosely. It seems to be about equivalent to allele or phenotype derived from a particular allele. At least in genetic circles. Anyone who knows want to comment?
a "trait" clearly doesn't have to be an advantage or disadvantage it is just some particular inheritable characteristic of someone.
And,yes, one clearly can be born better for running in a number of ways. I think that would be at too gross a level to fit what I think "trait" means. Lance Armstrongs "trait" of long thighs which happen to be advantages for cycling might be what we mean by trait.
I think you are tangleing up too many different ideas under one heading. But then I'm making up this whole post out of thin air.