The folks that published previously on the greenish warblers have some new work out: Irwin, et al.,
Science, Vol 307, 414-416, 21 January 2005. It gets into genetic measures of how all the warblers in the ring differ, and find that, sure enough, the two northernmost subspecies differ the most, even though their ranges overlap.
From 105 greenish warblers at 26 sites throughout the breeding range we obtained 62 AFLP markers that were variable and could be scored unambiguously as present or absent in each individual. West Siberian viridanus and east Siberian plumbeitarsus are clearly separated in AFLP genotypes, which confirms that the two taxa are genetically distinct. In contrast, AFLP genotypes change gradually through the ring of populations to the south (Fig. 1).