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Author Topic:   Evolution and Increased Diversity
New Cat's Eye
Inactive Member


Message 114 of 140 (440090)
12-11-2007 2:07 PM
Reply to: Message 113 by Elmer
12-11-2007 12:08 PM


The Dice Analogy
Hey Elmer,
There are two replies to your post. This one about the dice-stuff and the other one about the non-dice-stuff. I think the other one is more important so skip to that one if you are time constrained.
But let us not lose sight of the fact that we are discussing 'natural selection' in terms of biodiversity, i.e . Therefore any valid analogy between dice and the biodiversity of the globe or of any given ecosystem must be directly related to a comparison of 'biodiversity' to a 'die'.
Not necessarily. Each number on a die represents a genotype. Each die is an individual in the population, where the population is all the dice. Biodiversity is the number of different genotypes, or the amount of different numbers (1-6) on the dice.
The first thing we notice is the assumption that if biodiversity is analogous to a die, then biodiversity can never increase. Why not? Because with a 'die' what you start out with is a numbered six-sided object, a cube. A cube will always have no more, and no less, than six sides, no matter how much you toss it around. And a die, if it is to remain a die, will always be numbered 1-6. No side will ever turn blank, and no side will ever show a '7' or a '0'.
But that is not how the analogy has been defined.
Therefore the analogy of biodiversity to a die used in some sort of iterated 'rolling' process is false and invalid
Sure, your strawman analogy has been thoroughly refuted.
Now, lets move on to my analogy.
Now, if you try to claim that biodiversity is not analogous to a single 'die', but rather, to a given quantity of geometrically and numerically identical dice, then I am forced to point out to you that where all members of a given set are identical, [no matter how large or how small the sum of that set's members may be], then there is, by definition, no diversity in that set.
That’s correct. But in my analogy, before all the dice become number 1’s, I change the “rules”, which corresponds to the environment changing. Therefore, before the biodiversity reaches a value of zero (when all the dice are 1’s), and after the environment changes, we can actually have an increase in biodiversity as we will have both 1’s and 2’s (per the new rule), as well as the other non-6 numbers that have not been re-rolled yet.
So if that set is said to represent 'biodiversity', then biodiversity equals zero. And if that set consists of 1,000 identical dice, or only a single die, or any quantity in between, there will still be, only and always, six-sided cubes with the numbers 1-6 attached to their sides.
You don’t need the dice to change to represent a change in biodiversity. The numbers on the dice are the genotypes, so if the number of genotypes changes (for example if it went from eighty 1s, and twenty 3’s to eighty 1’s, ten 2’s and ten 3’s, then the biodiversity has increased from two genotypes to three), then we have had an increase in biodiversity.
One of the problems is that you will actually need to throw extra die in on each re-roll so that the population doesn’t dwindle and the analogy does a better job representing reality. The original analogy does need to be changed as exposed by MartinV.
But you seem to want to relate biodiversity, [as is the common practice among darwinians], not the die or the dice themselves, but rather the numerals, 1-6, afixed to their sides. Leaving aside the issues of analogizing biodiversity,-- that which involves concrete entities [bioforms],-- to that which is immaterial, incorporeal, ideational, and symbolic [numbers]
Yeah, that’s pretty much what I want. And I would expect that an analogy would require that something has been analogized , I mean, duh.
I must point out that a/ subtracting a particular numeral when it does not show face-up is to decrease the diversity showing 'across the board', and so, for that matter' is the act of 'keeping' only 6's, or any other number.
Yes, after a few generations, the biodiversity will decrease per the “rules” of the game. It is after the rules change (ie a change in the environment), that we will see an increase in biodiversity. [this is not to say that and increase in biodiversity requires a change in the environment, that is just the case for this analogy].
All you do in your analogy is a/ decrease the number of dice in play,
Yeah, that was an error (pointed out by MartinV). We will need to add dice back into the poor for the analogy to work more properly. Sorta replenish the pool, if you will.
and b/, decrease the variation in numerals that 'turn up', until you end with only identical numerals, perhaps only a single die, perhaps 50, no matter, all exhibiting the same identical numeral. In short, your analogy shows that biodiversity can fluctuate, at least among symbolic representations, by means of 'selection', but only in a negative, subtractive, decreasing sense.
Right, until the rules change and we start keeping both 1’s and 2’s. Then we can have an increase in biodiversity. It helps that we are replacing the removed dice now, so that if the popualtion becomes all 1’s, we can still have the increase. The original idea, was that the rules would be changed before all the dice became 1’s.
That is, no matter how many 1's you subtract, nor how many 6's you keep, your 'biodiversity' will never exceed in quantity the _six_ variables with which you began, but _must_ always end with a decreased number of variables.
I don’t think it matters how many variables we began with. Here’s why. After a few re-rolls (generations), we will have mostly 1’s with a few stragglers of the other numbers and no 6’s. Lets say we did not roll any 2’s the last time, so what we have is biodiversity value of four. Four numbers are present, 1,3, 4, and 5. If we change the rules to now keep the 6’s, or the 2’s or whatever and then we re-roll. We could then have five numbers present, which would be an increase in biodiversity.
All the way down to 1 numeral only, which, even if it shows multiple times, is still invariable, and thus the diversity shown is zero.
Yeah, I’ve corrected for that, it was an error on my part. “My bad.”
But you know, now that I’ve typed all this out, I remember why I don’t like analogies that much. More time is spent explaining the analogy than making the actual point of the analogy.
This analogy isn’t working out all that great, we can just drop it if you want. Or if you want to explore it further, I can do that too.
As far as the rest of your message, I’ll reply to that in the following post.

This message is a reply to:
 Message 113 by Elmer, posted 12-11-2007 12:08 PM Elmer has not replied

  
New Cat's Eye
Inactive Member


Message 115 of 140 (440092)
12-11-2007 2:09 PM
Reply to: Message 113 by Elmer
12-11-2007 12:08 PM


IOW, your 'natural selection' can reduce a given amount of biodiversity all the way to zero, but it can never increase it by so much as a fraction.
Hey alright! A contention! And it is the entire point of the thread. Hot damn!
I believe that natural selection, acting on random mutations, can lead to an increase in biodiversity.
It was this flaw that, when pointed out to Darwin, sent him back to Lamarck [unacknowledged] for his 'pangenesis' explanation for increased variation and added complexity, i.e., increased biodiversity, among bioforms. And it is what prompted Fisher, Haldane, and Wright to postulate 'random genetic mutation' as the source of _increased_ biodiversity, simply because 'natural selection' could not and did not account for it.
I don’t have a problem with adjusting/fixing scientific theories, as errors are found, do you?
I agree that NS cannot account for it alone. You need the source of the change in the genome in order to get the diversity. RM fits the bill.
If the environment is able to support more species and provide more niches for them, then we would expect, as more and more mutations arise,
Don't you see? You are now attributing increased biodiversity to random [supposedly] genetic mutation, and have entirely dropped 'ns' as the source/origin of novelty, i.e., increased biodiversity.
My position has been the same the whole time. RM provides for the source of the new information needed to allow for a change in the bioforms (what I was calling the “opportunity”), and NS is needed to ”decide’ which changes will be kept and which will be rejected. Neither one of them alone accounts for the increase in biodiversity, they have to work together to get it (except for the rare case of genetic drift, but I doubt that would lead to speciation so we can just forget about it).
After your 'random genetic mutation' supposedly increases the amount biodiversity present, your 'ns' , [whatever it is supposed to be, empirically-speaking(?!?)], merely limits, constrains, and reduces the amount of biodiversity that persists over time.
No, if a trait is advantageous it will become more prominent. It has been selected ”for’. Also, NS can remove all traits that are not the one selected for. Oh wait, is that what you mean? That you can only select ”for’ by removing the ”others’?
your 'ns' , [whatever it is supposed to be, empirically-speaking(?!?)]
Here is a link the wiki article on NS.
When you say 'selected for', you do realise that you are in fact saying is, "will not be eliminated by 'natural selection'", don't you?
Okay, now I see the angle you are coming from.
So now you are saying that 'random genetic mutation' is -not- the source of novel variation and increased complexity, i.e., the originating mechanism of increased biodiversity, but rather, that the 'environment is the generator of novelty and diversity.
Okay, okay. I think I see the problem.
RM is the “source” of the information that NS selects against. It provides the “opportunity” for an increase in biodiversity. NS is the factor that “decides” which traits are promoted (through the elimination of all others). Neither one of them alone can account for the increase. It is when they are combined that we can see an increase in biodiversity.
So basically the question boils down to: How can biodiversity increase when the selective factor can only removing unacceptable traits?
You seen to agree that RM can provide the new info to select from, but your contention seems to lie with the idea that by removing traits we can have an increase in them?
Let me ask you if that correctly reflects your position before we move forward and I respond to it, okay?



Why not? Is it possible that it is by some reason that you simply do not know?
No, it is because of what I _do know_ about 'natural selection'. See above. There is nothing that I do not know about it.
Holy shit! I guess I’m wasting my time then, huh? For some reason, I doubt it
Without wishing to insult you, anyone who believes that 'natural selection' can increase biodiversity simply does not understand 'natural selection', and/or the concept of biodiversity.
Don’t worry about insulting me, this is the internet. And I’m more than willing to admit that the ToE’s NS is erroneous and cannot account for the increase in biodiversity, but you have not convinced me as such, yet.
I don’t think the mutation is the “cause”. It just provides the “opportunity”.
Your fellow-darwinians would, I think, disagree with you and say that random chance, i.e., genetic accidents, are the "cause", and that it is the "environment" that "just provides the opportunity".
Isn’t it possible that we (me and my fellow “darwinians” [BTW, what is a ”darwinian’?]) are saying the same thing (that we mean the same thing) but I am just using the terms differently?
They will tell you that the idea of 'the environment' generating novely [eg., air generating birds,or deserts generating camels, either via morphological or genetic mutation], is 'lamarckian', [even though that is a distorted strawman of actual lamarckian theory].
But that is not what I mean by the environment “causing” it. The cause of the increase in biodiversity is in the selecting of the different mutations. The mutations, themselves, however, are what cause the change in the properties of what is being selected from.
You’ve conflated uses of the word “cause”. RM ”causes’ the change in the individual, NS ”causes’ the change in the population. But you can’t change the population with changes in the individuals. So now you’re saying that I’m saying that what changes the population is what changes the individual, but that is not what I am saying.
So far, I have shown that "NS" has no contribution to make to-increased- biodiversity, even though, theoretically, it may be said to account for -decreased- biodiversity. That leaves only "Random Genetic Mutation" to account for increased biodiversity, at least in terms of, 'modern evolutionary biology', current neo-darwinism is commonly designated.
RM is the ”source’ of the information that NS can select from, by the removal of unwanted traits, to provide an increase in biodiversity.
How that is possible is what I intend to explain to you. However, I have little time left right now.
That is the explanation you’re looking for, right?
We will cover that in my next post.
Let me know if you wish me to continue.
Yes, please do.

This message is a reply to:
 Message 113 by Elmer, posted 12-11-2007 12:08 PM Elmer has replied

Replies to this message:
 Message 117 by Elmer, posted 12-11-2007 8:14 PM New Cat's Eye has replied

  
New Cat's Eye
Inactive Member


Message 120 of 140 (440267)
12-12-2007 11:56 AM
Reply to: Message 117 by Elmer
12-11-2007 8:14 PM


RM is the ”source’ of the information that NS can select from, by the removal of unwanted traits, to provide an increase in biodiversity.
How that is possible is what I intend to explain to you. However, I have little time left right now.
Well, convincing me that subtraction equals addition is, I suspect, going to be one heck of a hard job; but you are free to try it on. Good luck.
“Subtraction equals addition” (although worded poorly) when your influx is greater than your retractions. Where ”influx’ is the alleles provided by RM and a ”retraction’ is selecting against an allele via NS.
What I have said is that you are confusing increased biodiversity [an increase in the number of taxa ] with population expansion [an increase in the number of individuals within the same taxon].
I guess I can see how you might think that, but I am not confusing those things.
You seen to agree that RM can provide the new info to select from, but your contention seems to lie with the idea that by removing traits we can have an increase in them?
Uhm, yes, I do distinguish between addition and subtraction, and insist that the one cannot be the other.
Okay smartass I wanted to make sure you weren’t in the “RM cannot add info” crowd.
So basically the question boils down to: How can biodiversity increase when the selective factor can only removing unacceptable traits?
I think it would be better phrased this way:--How can biodiversity increase where the causal factor to which the increase is attributed [ns] is only capable of removing traits from the total number of presently existing biodiverse traits.
You have to have more alleles being introduced to the population, by RM, than there are being removed from the population, by NS.
However, Wright and Kimura, as well as a number of 'anti-adaptationists', insist that 'rgm', all by itself, without any 'ns' at all, can and does increase biodiversity.
I would call that Genetic Drift. It is possible for it to lead to an increase in biodiversity but improbable.
By 'prominent' I suppose you to mean 'numerically greater relative to other traits'. But how do you know that it is 'advantageous'? Because it has become 'numerically greater relative to other traits'?!? This is the old 'fitness' tautology. Like all tautologies and circular arguments, it sounds great,-- but means nothing.
An allele is advantageous when it affects the phenotype in a way that increases the chance of the allele to be present in future generations in higher quantities.
Actually, I mean that 'to select for' is meaningless. "Selection", in the notional darwinian sense of 'natural selection', can only 'subtract from', never 'add to', the biodiversity available to it. So, as I've said again and again, where increased biodiversity is concerned, 'positive selection' aka 'positive natural selection', is merely a fantasy.
The selection is positive when it is not selected against. What would you call a number that is non-zero and non-negative? I see what you mean that NS isn’t really “adding” anything to the biosphere, but if it is non-zero and non-negative, then calling it positive seems fine to me. Unless, you want to turn this into a semantics argument? (I don’t.)
You get a constant influx of mutations to the population from RM. In a simplified model, NS just puts pressure on some of those mutations so that it is harder for them to reproduce. However, NS can also promote the reproduction of that mutation, if the mutation provides a benefit to reproducing.
Now, we can imagine mutations coming in from all over the place, in a way that each individual is more like a new bioform that it is a member of the parent bioform. In this case, every bioform that did not get selected against, or had a benefit that made reproduction easier, would be an increase in the biodiversity. In this case we would see biodiversity increase readily and quickly.
This, however, is not how it works in the real world. What we do have is a relatively small amount of mutations and those mutation are such that the new bioform is hardly distinguishable from the parent, if at all. It works out in a way that one allele might have an 89.67% chance of being passed on, and another has a 90% chance. This slight difference in that chance, when compounded over many generations leads to significant changes in the bioform. As long as there are significant changes in the bioform, speciation events are possible, and biodiversity can increase.
It is NS that provided that extra third of a percent chance of the allele being passed on. That is the positive selection that can “cause” the increase in biodiversity.
But on the semantic side, it doesn’t really “cause” it because the new information has been provided by RM, not NS. So, we can say that RM causes it, but without the selective pressure, the chances of being passed on would be random, and it would boil down to the “its just so” case you’ve mentioned. It would be totally stochastic. But that is not what we observe.
The point is, that if 'rgm' can generate novel variations on its own, but 'ns'cannot generate novelty off its own bat, then 'rgm' may theoretically proposed as the source of increased biodiversity, but 'ns' cannot be.
Yes, NS is not the ”source’. But it can and does provide non-zero, non-negative pressure on bioforms so that their genotype has an actual better chance of reproduction, which causes the variation within the population, which is what can lead to a speciation event and thus an increase in biodiversity.
By which you mean that it has not been 'selected against', meaning only that it was not wiped out, eradicated, or even diminished numerically, by 'ns'. 'Not killing' someone is not the same as bringing new life into the world. Sparing a life is not saving a life.
But from RM, we have a constant addition of new “lives”. To not remove a new life means that bringing new life into the world is the default.
Environmental factors result in the differential mortality rates deaths of different organisms. Some organisms die sooner than others, from an almost infinite variety of particular causes, most of them environmental and/or organismic. Dressing up this fact/effect with labels like 'negative selection' and 'positive selection', and then awarding causal power to a label/fact/effect, is inane.
What about it, specifically, makes it inane?
The causal power is awarded because NS can have an effect that makes the allele more beneficial, that the allele has a higher probability of succeeding. RM alone doesn’t provide this benefit. It takes that environment to put the pressure on all the others, or remove pressure from the one in order for the diversity to increase.
Besides, as I've already told you, having more individuals with the same bioform does not increase biodiversity--only having more varieties of bioform does that.
The increase in individuals with the allele are technically in the same bioform, but as more and more mutations stack up, the individuals become more and more distinct from that bioform and when a speciation event occurs, the biodiversity increases.
The biodiversity of the great plains did not change just because there were more or fewer bison grazing on it. It changed when new bioforms [the horse, domestic animals, etc.] were added to it, or, [like the passenger pigeon and others extinct lifeforms], were subtracted from it.
But speciation does not work like: bison --> horse.
It’d be more like: bison --> bisoe --> bosoe --> bosse --> hosse --> horse.
Where the bisoe split from the bison, while the bison remained. The bisoe and bison, are not easily recognized as different bioforms, just like the bosoe and the bisoe aren’t, it is only after much change that we can recognize that a horse is no longer in the same bioform as a bison.
Meaning that 'rgm' supposedly causes the change in bioform, [additional bioform being an increase in biodiversity], and 'ns' supposedly causes a change in the numbers of the same bioform [no increase in biodiversity].
That seems about right. And that’s where the conflation of the word “cause” comes into play. Can we say that either one of them really causes biodiversity, can’t we say that they both cause it? It depends on how we use the word “cause”.
I don’t have a problem with adjusting/fixing scientific theories, as errors are found, do you?
No, I certainly do not. But it is you, (not I, and not Darwin, nor Fisher, et al), that continues to insist that 'natural selection' can and does increase biodiversity. Which indicates that, as a matter of fact, you do "have a problem with adjusting/fixing scientific theories, as errors are found".
What do you propose is the “fix” to the ToE that I would have a problem with?
Just for shits and giggle, what if I concede that NS cannot do it, then what do you propose IS the “cause” of the increase in biodiversity that we observe?

This message is a reply to:
 Message 117 by Elmer, posted 12-11-2007 8:14 PM Elmer has replied

Replies to this message:
 Message 121 by RAZD, posted 12-12-2007 12:54 PM New Cat's Eye has not replied
 Message 122 by Elmer, posted 12-12-2007 4:11 PM New Cat's Eye has not replied

  
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